Thursday, February 26, 2009

Sloths

Sloth
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This article is about the South American mammal. For other uses, see Sloth (disambiguation).
Sloths [1]


Brown-throated Three-toed Sloth
(Bradypus variegatus)
Gatun Lake, Republic of Panama.
Scientific classification
Kingdom: Animalia

Phylum: Chordata

Class: Mammalia

Subclass: Theria

Infraclass: Eutheria

Superorder: Xenarthra

Order: Pilosa

Suborder: Folivora
Delsuc, Catzeflis, Stanhope, and Douzery, 2001

Families
Bradypodidae
Megalonychidae
†Megatheriidae
†Mylodontidae
†Orophodontidae
†Nothrotheriidae

The living sloths comprise six species of medium-sized mammals that live in Central and South America belonging to the families Megalonychidae and Bradypodidae, part of the order Pilosa. The sloth's taxonomic suborder is Folivora, while some call it Phyllophaga. Both names mean "leaf-eaters"; the first is derived from Latin, the second from ancient Greek. Tribal names include Ritto, Rit and Ridette, mostly forms of the word "sleep", "eat" and "dirty" from Tagaeri tribe of Huaorani.

Contents [hide]
1 Megatherium
2 Ecology
3 Physiology
4 Classification of the sloth
5 Gallery
6 References
7 External links



Megatherium
Until geologically recent times, large ground sloths such as Megatherium[2] lived in South America and parts of North America, but along with many other animals they disappeared immediately after the arrival of humans on the continent[3] Much evidence suggests that human hunting contributed to the extinction of the American megafauna, like that of far northern Asia, Australia, New Zealand, and Madagascar [4]. Simultaneous climate change that came with the end of the last Ice age may have also played a role in some cases. However, the fact that ground sloths survived on the Antilles long after they had died out on the mainland points towards human activities as the agency of extinction.


Ecology
The living sloths are omnivores. They may eat insects, small lizards, and carrion, but their diet consists mostly of buds, tender shoots, and leaves, mainly of Cecropia trees. They have made extraordinary adaptations to an arboreal browsing lifestyle. Leaves, their main food source, provide very little energy or nutrition and do not digest easily. Sloths therefore have very large, specialized, slow-acting stomachs with multiple compartments in which symbiotic bacteria break down the tough leaves. As much as two-thirds of a well-fed sloth's body-weight consists of the contents of its stomach, and the digestive process can take a month or more to complete.

Even so, leaves provide little energy, and sloths deal with this by a range of economy measures: they have very low metabolic rates (less than half of that expected for a creature of their size), and maintain low body temperatures when active (30 to 34 °C or 86 to 93 °F), and still lower temperatures when resting.

Although unable to survive outside the tropical rainforests of South and Central America, within that environment sloths are outstandingly successful creatures: they can account for as much as half the total energy consumption and two-thirds of the total terrestrial mammalian biomass in some areas.[citation needed] Of the six living species, only one, the Maned Three-toed Sloth (Bradypus torquatus), has a classification of "endangered" at present. The ongoing destruction of South America's forests, however, may soon prove a threat to other sloth species.


Physiology

Hoffmann's Two-toed Sloth, Choloepus hoffmanni.Sloth fur exhibits specialized functions: the outer hairs grow in a direction opposite from that of other mammals. In most mammals, hairs grow toward the extremities, but because sloths spend so much time with their legs above their bodies, their hairs grow away from the extremities in order to provide protection from the elements while the sloth hangs upside down. In moist conditions, the fur hosts two species of symbiotic cyanobacteria, which provide camouflage[5][6]. Because of this algae, sloth fur is a small ecosystem of its own, hosting many species of non-parasitic insects.[7]. Sloths have short, flat heads; big eyes; a short snout; long legs; and tiny ears. They also have stubby tails, usually 6-7cm long. Altogether, sloths' bodies usually are anywhere between 50 and 60 cm long.

Sloths' claws serve as their only natural defense. A cornered sloth may swipe at its attackers in an effort to scare them away or wound them. Despite sloths' apparent defenselessness, predators do not pose special problems: sloths blend in with the trees and, moving only slowly, do not attract attention. Only during their infrequent visits to ground level do they become vulnerable. The main predators of sloths are the jaguar, the harpy eagle, and humans. The majority of sloth deaths in Costa Rica are due to contact with electrical line and poachers. Despite their adaptation to living in trees, sloths make competent swimmers. Their claws also provide a further unexpected deterrent to human hunters - when hanging upside-down in a tree they are held in place by the claws themselves and often do not fall down even if shot from below.


Pale-throated Three-toed Sloth (Bradypus tridactylus) in a Costa Rican rehabilitation center.Sloths move only when necessary and even then very slowly: they have about a quarter as much muscle tissue as other animals of similar weight. They can move at a marginally higher speed if they are in immediate danger from a predator (4.5 m or 15 feet per minute), but they burn large amounts of energy doing so. Their specialized hands and feet have long, curved claws to allow them to hang upside-down from branches without effort[8]. While they sometimes sit on top of branches, they usually eat, sleep, and even give birth hanging from limbs. They sometimes remain hanging from branches after death. On the ground their maximum speed is 0.54 meters per second (1.2 mph)[9].

It had been thought that sloths were among the most somnolent animals, sleeping from 15 to 18 hours each day. Recently, however, Dr. Neil Rattenborg and his colleagues from the Max Planck Institute for Ornithology in Starnberg, Germany, published a study testing sloth sleep-patterns in the wild; this is the first study of its kind. The study indicated that sloths sleep just under 10 hours a day.[10] They go to the ground to urinate and defecate about once a week. They go to the same spot each time and are vulnerable while doing so. The reason for this risky behavior is unknown, although some believe that this is to avoid making noise while defecating from up high that would attract predators.[11]

Infant sloths normally cling to their mother's fur, but occasionally fall off. Sloths are very sturdily built and rarely die from a fall. In some cases they die from a fall indirectly because the mothers prove unwilling to leave the safety of the trees to retrieve the young. Females normally bear one baby every year, but sometimes sloths' low level of movement actually keeps females from finding males for longer than one year.

Almost all mammals have seven cervical vertebrae or "neck bones" (including those with very short necks, such as elephants or whales, and those with very long necks, such as giraffes). The few exceptions include manatees and two-toed sloths, which each have only six cervical vertebrae, and three-toed sloths with nine cervical vertebrae.[12]


Classification of the sloth
The living sloths belong to one of two families, known as the Megalonychidae ("two-toed" sloths) and the Bradypodidae (three-toed sloths). All living sloths have in fact three toes; the "two-toed" sloths, however, have only two fingers. Two-toed sloths are generally faster moving than three-toed sloths. Both types tend to occupy the same forests: in most areas, one species of three-toed sloth and one species of the larger two-toed type will jointly predominate.

However, their adaptations belie the actual relationships of the living sloth genera, which are more distant from each other than their outward similarity suggests. The common ancestor of the two genera apparently lived 35-40 million years ago, making the living forms stunning examples of convergent or parallel evolution.[13] The two-toed sloths of today are far more closely related to one particular group of ground sloths than to the living three-toed sloths. Whether these ground-dwelling Megalonychidae were descended from tree-climbing ancestors or whether the two-toed sloths are really miniature ground sloths converted (or reverted) to arboreal life cannot presently be determined to satisfaction. The latter possibility seems slightly more likely, given the fact that the small ground sloths Acratocnus and Neocnus which were also able to climb are among the closer relatives of the two-toed sloths, and that these together were related to the huge ground sloths Megalonyx and Megalocnus.

The evolutionary history of the three-toed sloths is not at all well-known. No particularly close relatives, ground-dwelling or not, have yet been identified.

The ground sloths do not constitute a monophyletic group. Rather, they make up a number of lineages, and as far as is known until the Holocene most sloths were in fact ground-dwellers. The famous Megatherium for example belonged to a lineage of ground sloths that was not very close to the living sloths and their ground-living relatives like the small Neocnus or the massive Megalonyx. Meanwhile, Mylodon, among the last ground sloths to disappear, was only very distantly related to either of these.

ORDER PILOSA[1]
Suborder Folivora
Family Bradypodidae
Genus Bradypus (Three-toed sloths)
Pygmy Three-toed Sloth, Bradypus pygmaeus
Maned Three-toed Sloth, Bradypus torquatus
Pale-throated Three-toed Sloth, Bradypus tridactylus
Brown-throated Three-toed Sloth, Bradypus variegatus
Family Megalonychidae (includes extinct megalonychid ground sloths)
Genus Choloepus (Two-toed sloths)
Linnaeus's Two-toed Sloth, Choloepus didactylus
Hoffmann's Two-toed Sloth, Choloepus hoffmanni
Family †Megatheriidae: megatheriid ground sloths
Family †Mylodontidae: mylodontid ground sloths
Family †Orophodontidae: orophodontid ground sloths
Family †Nothrotheriidae: nothrotheriid ground sloths

Suborder Vermilingua (anteaters and tamanduas)

Gallery

Brown-throated Three-toed Sloth (Bradypus variegatus), Cahuita National Park, Costa Rica.

Choloepus sp., Dortmund Zoo.

Megatherium fossil.

Scelidotherium leptocephalum fossil. Muséum national d'histoire naturelle, Paris.


Paramylodon harlani fossil, National Museum of Natural History, Washington, D.C.

Brown-throated Three-toed Sloth The only publicly displayed Three-toed in the US. Dallas World Aquarium and Zoo, Dallas, TX.

Reconstruction of giant ground sloth based on Megalonyx jeffersonii, Iowa Museum of Natural History.



References
^ a b Gardner, Alfred (November 16, 2005). Wilson, D. E., and Reeder, D. M. (eds). ed. Mammal Species of the World (3rd edition ed.). Johns Hopkins University Press. pp. 100-101. ISBN 0-801-88221-4. http://www.bucknell.edu/msw3/browse.asp?id=11800002.
^ Wildfacts "Megatherium". BBC Science & Nature. http://www.bbc.co.uk/nature/wildfacts/factfiles/456.shtml Wildfacts. Retrieved on 2007-01-09.
^ Wildfacts "Megatherium". BBC Science & Nature. http://www.bbc.co.uk/nature/wildfacts/factfiles/456.shtml Wildfacts. Retrieved on 2007-01-09.
^ Wildfacts "Megatherium". BBC Science & Nature. http://www.bbc.co.uk/nature/wildfacts/factfiles/456.shtml Wildfacts. Retrieved on 2007-01-09.
^ Rainforest Canopy—Animals
^ The Hidden Lives of Sloths: Interesting Thing of the Day
^ Minds in the Making | Science and Technology | Wisdom in Nature
^ Mendel, 1979; 1981a; 1981b; 1985
^ http://hypertextbook.com/facts/1999/IvyWeinberg.shtml
^ http://news.bbc.co.uk/2/hi/science/nature/7396356.stm BBC News Article "Sloth's Lazy Image 'A Myth'"
^ David Attenborough, Life of Mammals, BBC
^ Narita, Yuichi; Kuratani, Shigeru (2005), "Evolution of the Vertebral Formulae in Mammals: A Perspective on Developmental Constraints", Journal of Experimental Zoology (Mol Dev Evol) 304 (2): 91–106, doi:10.1002/jez.b.21029
^ Gaudin, Timothy J. (2004-02-03). "Phylogenetic relationships among sloths (Mammalia, Xenarthra, Tardigrada): the craniodental evidence". Zoological Journal of the Linnean Society (Linnean Society of London) 140 (2): 255–305. doi:10.1111/j.1096-3642.2003.00100.x. http://www3.interscience.wiley.com/journal/118759635/abstract?CRETRY=1&SRETRY=0. Retrieved on 2008-12-16.

External links
Look up sloth in
Wiktionary, the free dictionary.
Wikimedia Commons has media related to: Folivora
Two-toed Sloth Page at National Geographic website
Three-toed Sloth Page at National Geographic website
Caltech Sloth Page
Aviarios del Caribe Sloth Sanctuary (open to tourists, and close to the cruise ship pier, in Costa Rica).
Sloth World: An online bibliography and database of sloth papers from around the world
Pictures from sloths.org
Movie depicting baby sloth
A website about sloths and the rain forest
[show]v • d • eExtant Pilosa species by suborder

Kingdom Animalia · Phylum Chordata · Class Mammalia · Infraclass Eutheria · Superorder Xenarthra

Folivora (Sloths)


Bradypodidae Bradypus
(Three-toed sloths) Pygmy Three-toed Sloth (B. pygmaeus) · Maned Sloth (B. torquatus) · Pale-throated Sloth (B. tridactylus) · Brown-throated Sloth (B. variegatus)


Megalonychidae Choloepus
(Two-toed sloths) Linnaeus's Two-toed Sloth (C. didactylus) · Hoffmann's Two-toed Sloth (C. hoffmanni)


Vermilingua (Anteaters)


Cyclopedidae Cyclopes Silky Anteater (C. didactylus)


Myrmecophagidae Myrmecophaga Giant Anteater (M. tridactyla)

Tamandua
(Tamanduas) Northern Tamandua (T. mexicana) · Southern Tamandua (T. tetradactyla)


Category


Retrieved from "http://en.wikipedia.org/wiki/Sloth"
Category: Pilosans
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